Depressing the Mode button shifts the transmission into (or out of) Economy mode, which changes the shift points by approx 200 RPM.
Most coaches have their transmission pre-programmed to Normal or Standard mode, and the driver can choose to select Economy mode. Some coaches, such as Marathon Prevost conversions, have their transmission pre-programmed to Economy mode, and the driver can choose to select Normal mode. According to Allison, many buyers of Marathons have asked to have their transmissions re-programmed to Normal/Standard mode.